Ed Rajotte, Ph.D.

  • Professor of Entomology and IPM Coordinator
Ed Rajotte, Ph.D.
508 Ag Sciences & Industries Building
University Park, PA 16802
Work Phone: 814-863-4641

Connect with Ed Rajotte, Ph.D.



  1. B.S., Biology, University of Connecticut, 1973
  2. M.S., Entomology, University of Connecticut, 1975
  3. Ph.D., Entomology, Rutgers University, 1981

Current Graduate Students:


Sulav Paudel

Asifa Hameed

Ngoc Phan

Mariam Taleb

Recent Grads:


Alexis Barbarin

Naworaj Acharya

Sarah Shugrue

Mitzy Porras

Megan Wilkerson

Maria Mazin

Department Focus Areas:

Ecological Applications

International Development


IPM, international agriculture, pest management policy, biology of fruit pests, pollination and apiculture (


Integrated Pest Management, Comparative European and U.S. Agriculture


Integrated pest management; international agriculture; IPM in schools, urban IPM programming.

Research Activities & Interests :

My research focuses on insect ecology and management in support of integrated pest management systems. Domestically, I concentrate on biological, social and economic issues in integrated pest management, bee management and pollination, pest management in under-served urban neighborhoods and fruit IPM. Internationally, I have had projects in South Asia, Southeast Asia, Eastern Europe and North Africa primarily designing and testing IPM approaches in tropical vegetables, IPM information systems and impact analysis.

Extension Activities & Interests:

As IPM Coordinator for Pennsylvania Cooperative Extension, I represent and promote IPM with federal, state and municipal governments; represent Pennsylvania to state and federal policy-making bodies on pest management issues; coordinate IPM in schools and urban environments.  Our extension IPM programs are team efforts involving close cooperation among members of the entomology department and extension faculty and staff in other departments and in the counties. The Pennsylvania IPM Program represents Penn State's disciplines including entomology, plant pathology, weed science, nematology and others.

Recent Publications:

(See also:

Sulav Paudel, Po-An Lin, Majid R. Foolad, Jared G. Ali, Edwin G. Rajotte, Gary W. Felton: Induced Plant Defenses Against Herbivory in Cultivated and Wild Tomato. Journal of Chemical Ecology 08/2019;, DOI:10.1007/s10886-019-01090-4

M.S. Hossain, B.C. Sarkar, M.M. Hossain, M.Y. Mian, E.G. Rajotte, R. Muniappan, M.E. O'Rourke: Comparison of biorational management approaches against mango fruit fly (Bactrocera dorsalis Hendel) in Bangladesh. Crop Protection 05/2019;, DOI:10.1016/j.cropro.2019.05.001

Maria Mazin, Stefanos S. Andreadis, Nina E. Jenkins, Kevin R. Cloonan, T.C. Baker, Ed G. Rajotte: Activity and distribution of the mushroom phorid fly, Megaselia halterata , in and around commercial mushroom farms. Entomologia Experimentalis et Applicata 04/2019;, DOI:10.1111/eea.12777

Joseph Walls, Edwin Rajotte, Cristina Rosa: The Past, Present, and Future of Barley Yellow Dwarf Management. 01/2019; Agriculture 9(1):23., DOI:10.3390/agriculture9010023

Biddinger, D., E. G. Rajotte, N. K. Joshi*.  2018.  Chapter. 4 Integrating pollinator health into tree fruit IPM- A case study of Pennsylvania apple production (Chapter- 4), p. 69-83.  In: The pollination of cultivated plants: a compendium for practitioners. Vol. 1. (Editor- D. Roubik). Food and Agricultural Organization of the United Nations. 313 p.

JF Martinez, C Rosa, C Rotella, M Terrones, E Rajotte. 2018. Application of a carbon-nanotube filter that traps and concentrates virions to improve the limits of detection of Tomato spotted wilt virus in tomato. PHYTOPATHOLOGY 108 (10), 62-63

Mitzy Porras, Consuelo M. De Moraes, Mark C. Mescher, Edwin G. Rajotte, Tomás A. Carlo: A plant virus (BYDV) promotes trophic facilitation in aphids on wheat. Scientific Reports 12/2018; 8(1)., DOI:10.1038/s41598-018-30023-6

Mia G. Park, Neelendra K. Joshi, Edwin G. Rajotte, David J. Biddinger, John E. Losey, Bryan N. Danforth: Apple grower pollination practices and perceptions of alternative pollinators in New York and Pennsylvania. Renewable Agriculture and Food Systems 04/2018;, DOI:10.1017/S1742170518000145

Lalit Sah, Y.Dhoj GC, Mukti Devkota, Luke A. Colavito, George W. Norton, Edwin George Rajotte, R. Muniappan, A. Singh: Evaluation of pest exclusion net to major insect pest of tomato in Kavre and Lalitpur. 2018. Nepalese Journal of Agricultural Science. 16, 135-144.

S Paudel, I Mikó, A Deans, E Rajotte, G Felton. 2018. Lyonet’s gland of the tomato fruitworm, Helicoverpa zea (Lepidoptera: Noctuidae). PeerJ Preprints 6, e26455v1

Maria Mazin, Stefanos S. Andreadis, Nina E. Jenkins, Edwin G. Rajotte: The mushroom sciarid fly, Lycoriella ingenua, benefits from its association with green mold disease (Trichoderma aggressivum) in commercial mushroom production. Journal of Pest Science 11/2017;, DOI:10.1007/s10340-017-0930-4

Neelendra K. Joshi, Edwin G. Rajotte, Kusum J. Naithani, Greg Krawczyk, Larry A. Hull: Population Dynamics and Flight Phenology Model of Codling Moth Differ between Commercial and Abandoned Apple Orchard Ecosystems. Frontiers in Physiology 09/2016; 7., DOI:10.3389/fphys.2016.00408

Walls, J.T., III, Piero Caciagli, John F. Tooker, Joseph M. Russo, Edwin G. Rajotte, Cristina Rosa: Modeling the decision process for barley yellow dwarf management. Computers and Electronics in Agriculture 09/2016; 127:775-786., DOI:10.1016/j.compag.2016.08.005

Melanie A Kammerer, David J Biddinger, Edwin G Rajotte, David A Mortensen: Local Plant Diversity Across Multiple Habitats Supports a Diverse Wild Bee Community in Pennsylvania Apple Orchards. Environmental Entomology 09/2015;  DOI:10.1093/ee/nvv147

D C Soergel, N Ostiguy, S J Fleischer, R R Troyer, E G Rajotte, G Krawczyk: Sunflower as a Potential Trap Crop of Halyomorpha halys (Hemiptera: Pentatomidae) in Pepper Fields. Environmental Entomology 09/2015;  DOI:10.1093/ee/nvv136

David J. Biddinger, Edwin G. Rajotte 2015. Integrated Pest and Pollinator Management – Adding a New Dimension To An Accepted Paradigm. Current Opinion in Insect Science. 06/2015; 10. DOI:10.1016/j.cois.2015.05.012

Joshi, Neelendra K. Timothy LeslieEdwin George RajotteMelanie KammererMark Otieno and David Biddinger 2015. Comparative Trapping Efficiency to Characterize Bee Abundance, Diversity and Community Composition in Apple Orchards. ANNALS OF THE ENTOMOLOGICAL SOCIETY OF AMERICA · MAY 2015. DOI: 10.1093/aesa/sav057.

Acharya, Naworaj, Edwin G. Rajotte, Nina E. Jenkins, Matthew B. Thomas. 2015. Influence of biotic and abiotic factors on the persistence of a Beauveria bassiana biopesticide in laboratory and high-rise poultry house settings. Biocontrol Science and Technology 06/2015; DOI:10.1080/09583157.2015.1055318 · 0.73 Impact Factor

Biddinger, David J., Edwin G. Rajotte. 2015. Integrated Pest and Pollinator Management – Adding a New Dimension To An Accepted Paradigm. Current Opinion in Insect Science; DOI:10.1016/j.cois.2015.05.012

Naworaj Acharya, Rebecca A. Seliga, Edwin G. Rajotte, Nina E. Jenkins, Matthew B. Thomas: Persistence and efficacy of a Beauveria bassiana biopesticide against the house fly, Musca domestica , on typical structural substrates of poultry houses. Biocontrol Science and Technology 01/2015; 25(6):1-41. DOI:10.1080/09583157.2015.1009872

Paudel, Sulav, Edwin G Rajotte, Gary W Felton. 2014.Benefits and costs of tomato seed treatment with plant defense elicitors for insect resistance. Arthropod-Plant Interactions 12/2014; 8(6):539-545. DOI:10.1007/s11829-014-9335-y 

Naworaj Acharya, Edwin G. Rajotte, Nina E. Jenkins, Matthew B. Thomas: Potential for biocontrol of house flies, Musca domestica, using fungal biopesticides. Biocontrol Science and Technology 11/2014; 25(5):513-524.

Naworaj Acharya, Edwin G. Rajotte, Nina E. Jenkins, Matthew B. Thomas: Potential for biocontrol of house flies, Musca domestica, using fungal biopesticides. Biocontrol Science and Technology 11/2014; 25(5). DOI:10.1080/09583157.2014.992862

Priscila Strapasson, Delia M Pinto-Zevallos, Sulav Paudel, Edwin G Rajotte, Gary W Felton, Paulo H G Zarbin: Enhancing Plant Resistance at the Seed Stage: Low Concentrations of Methyl Jasmonate Reduce the Performance of the Leaf Miner Tuta absoluta but do not Alter the Behavior of its Predator Chrysoperla externa.. Journal of Chemical Ecology 10/2014;  DOI:10.1007/s10886-014-0503-4

Alexis M. Barbarin, Corentin M. Barbu, Ron Gebhardtsbauer, Edwin G. Rajotte: Survival and Fecundity of Two Strains of Cimex lectularius (Hemiptera: Heteroptera). Journal of Medical Entomology 09/2014; 51(5). DOI:10.1603/ME12236

D. Biddinger, K. Demchak, E. Rajotte, T. Baugher, N. Joshi. 2014. Pollinators and Pesticide Sprays during Bloom in Fruit Plantings. Fruit Times. 04/2014

Alexis M. Barbarin, Corentin M. Barbu, Ron Gebhardtsbauer, Edwin G. Rajotte. 2014. Survival and Fecundity of Two Strains of Cimex lectularius (Hemiptera: Heteroptera). Journal of Medical Entomology 09/2014; 51(5)

Naworaj Acharya, Edwin G. Rajotte, Nina E. Jenkins, Matthew B. Thomas. 2014. Potential for biocontrol of house flies, Musca domestica, using fungal biopesticides. Biocontrol Science and Technology 11/2014

Priscila Strapasson, Delia M Pinto-Zevallos, Sulav Paudel, Edwin G Rajotte, Gary W Felton, Paulo H G ZarbinEnhancing Plant Resistance at the Seed Stage: Low Concentrations of Methyl Jasmonate Reduce the Performance of the Leaf Miner Tuta absoluta but do not Alter the Behavior of its Predator Chrysoperla externa. Journal of Chemical Ecology 10/2014

D. Biddinger, E. Rajotte, N. Joshi, A. Ritz. 2013. The Role of Pollen Bees in Fruit Tree Pollination and Some New Cautions on Pesticide Use. Fruit Times. 08/2013

G. S. Arida, B. S. Punzal, B. M. Shepard, Edwin George Rajotte. Parasitism of the leaf miner Lirionmyza trifolii (Burgess)(Diptera: Agromyzidae) larvae on onion grown after rice in Central Luzon, Philippines. The Philippine Entomologist. 10/2013; 27(2):100-108

Biddinger,David J,  Jacqueline L Robertson, Chris Mullin, James Frazier, Sara A Ashcraft, Edwin G Rajotte, Neelendra K Joshi, Mace Vaughn. 2013. Comparative Toxicities and Synergism of Apple Orchard Pesticides to Apis mellifera (L.) and Osmia cornifrons (Radoszkowski). PLoS ONE 09/2013; 8(9):1-6

Dinakaran, D. , G. Gajendran, S. Mohankumar, G. Karthikeyan, S. Thiruvudainambi, E. I. Jonathan, R. Samiyappan, D. G. Pfeiffer, E. G. Rajotte, G. W. Norton, S. Miller, R. Muniappan. 2013Evaluation of integrated pest and disease management module for shallots in Tamil Nadu, India - A farmer’s participatory approach. Journal of Integrated Pest Management. 07/2013; 4(2):B1-B9.

Biddinger, David J , Neelendra K Joshi, Edwin G Rajotte, Noemi O Halbrendt, Cassandra Pulig, Kusum J Naithani, Mace Vaughan 2013. An immunomarking method to determine the foraging patterns of Osmia cornifrons and resulting fruit set in a cherry orchard. Apidologie 07/2013; · 2.27 Impact Factor

Levitt, A.L., R.Singh, D.L. Cox-Foster, E. Rajotte, K. Hoover, N. Ostiguy and E.C. Holmes. 2013. Cross-species transmission of honey bee viruses in associated arthropods. Virus Research 07/2013; DOI:10.1016/j.virusres.2013.06.013

Sticky board traps for managing leaf miner, Liriomyza trifolii (Burgess) (Diptera: Agromyzidae) infestations in onion (Allium cepa Linn.)
G. S. Arida, B. S. Punzal, B. M. Shepard, Edwin George Rajotte. The Philippine Entomologist. 10/2013; 27(2):109-119

Barbarin, A.M., R. Gebhardtsbauer and E.G. Rajotte. 2013. Evaluation of blood regimen on the survival of Cimex lectularius L. using life table parameters. Insects 2013, 4(2), 273-286; doi:10.3390/insects4020273.

Barbarin, Alexis M., Nina E. Jenkins, Edwin G. Rajotte, Matthew B. Thomas, A preliminary evaluation of the potential of Beauveria bassiana for bed bug control, Journal of Invertebrate Pathology, Volume 111, Issue 1, 15 September 2012, Pages 82–85, ISSN 0022-2011, 10.1016/j.jip.2012.04.009

Geng, Jian, Zhi-Hong Li, Edwin G. Rajotte, Fang-Hao Wan, Xiao-Yu Lu, Zhi-Ling Wang. 2011. Potential geographical distribution of Rhagoletis pomonella (Diptera: Tephritidae) in China. Insect Science 18, 575-582.

Joshi N. K., L.A. Hull, E.G. Rajotte, G. Krawczyk, E. Bohnenblust. (2011) Evaluating sex pheromone and kairomone-based lures for monitoring of codling moth in apple orchards. Pest Management Science. (

Singh, Rajwinder, Abby L. Levitt, Edwin G. Rajotte, Edward C. Holmes, Nancy Ostiguy, Dennis vanEngelsdorp, W. Ian Lipkin, Claude W. dePamphilis, Amy L. Toth, Diana L. Cox-Foster. 2010. RNA Viruses in Hymenopteran Pollinators: Evidence of Inter-Taxa Virus Transmission via Pollen and Potential Impact on Non-Apis Hymenopteran Species. PLoS ONE; 5 (12): e14357

Zaman, F.U., D.D. Calvin, E.G. Rajotte and D.V. Sumerford. 2010. Can a specialist parasitoid, Macorcentrus cingulum, (Hymenoptera:Braconidae), influence the ecotype structure of its preferred host, Ostrinia nubilalis(Lepidoptera:Crambidae). J. Econ. Entomol. 103(2) 249-56. (

Arida, G.S., B.S. Punzal, and E.G. Rajotte. 2009. Effect of chlorpyrifos+BPMC insecticide spray on the population density, damage and natural enemies of leafminer (Lirioyza trifolii (Burgess)) on onion (Allium cepa Linn.) grown after rice (Oryza sativa Linn.), The Philippine Entomologist 23: 56-66.

Brewer, M.J., E.G. Rajotte, J.R.Kaplan, P.B. Goodell, D.J. Biddinger, J.N. Landis, R. M. Nowierski, B. F. Smallwood and M.E. Whalon. 2009. Opportunities, experiences and strategies to connect integrated pest management to U.S. Department of Agriculture conservation programs. American Entomologist 55(3) 140-46. (

Huang, H., Rajotte, E.G., Li, Z., Chen, K. and Zhang, S., 2009. A Web-Based Expert System for Assisted identification of Quarantine Stored Insect Pests. Information Processing, Volume 293, Computer and Computing Technologies in Agriculture II, Volume 1, eds. D. Li, Z. Chunjiang, (Boston: Springer), pp. 701–714.

Wang, N., Li, Z., Wu, J., Rajotte, E.G., Wan, F. and Wang, Z., 2009, The potential geographical distribution of Bactrocera dorsalis (DIPTERA: TEPHRIDIDAE) in China based on emergence rate models and ARCGIS. In IFIP International Federation for Information Processing, Volume 293, Computer and Computing Technologies in Agriculture II, Volume 1, eds. D. Li, Z. Chunjiang, (Boston: Springer), pp. 399–411.

Joshi  N. K; L A. Hull; G. Krawczyk; E. G. Rajotte (2008) Field results of mating disruption technologies for the control of codling moth, Cydia pomonella (L.), and oriental fruit moth, Grapholita molesta (Busck) in Pennsylvania apple orchards. Aspects of Applied Biology (84) 153-161.

Pilcher, C. and E.G. Rajotte. 2008. The role of assessment and evaluation in IPM implementation. In Integrated Pest Management. E. B. Radcliff and W. D. Hutchison [Eds]. Cambridge University Press. 

Joshi, N.K., Hull, L.A., Myers, C.T., Krawczyk, G., Rajotte, E.G. 2007. Oviposition preference of Oriental fruit moth [Grapholita molesta (Busck), Lepidoptera: Tortricidae] for apple cultivars. Proceedings of the 16th International Plant Protection Congress. 308-309.

Webster, N. and E.G. Rajotte. 2006. Service Learning: A model for integrating IPM methods and practices. Am. Entomol. 52(4) 218-220.

Martin, E.C., A.M. Baltazar, R.P. Robles, J.M. Ramos, E.G. Rajotte, L.T.Kok and S.K. DeDatta. 2005. Life Cycle and Efficacy of Hawaiian Beet Webworm (Spoladea recurvalis F.) as a Potential Biological Conrol Agent Against Horse Purslane (Trianthema portulacastrum L.). Philippine Journal of Weed Science (submitted).

Miller, S.A., A.M.N. R. Karim, A.M. Baltazar, E.G. Rajotte and G.W. Norton. Developing IPM Packages in Asia. P 27-50. In Norton, G. W., E.A. Heinrichs, G.C. Luther and M.E. Irwin (eds.) 2005. Globalizing Integrated Pest Management: A participatory research process. Blackwell Publishing, Ames Iowa. 338 pp.

Norton, G. W., E.G. Rajotte and G.C. Luther. The Paricipatory Integrated Pest Management (PIPM) Process. p 13 – 26. In Norton, G. W., E.A. Heinrichs, G.C. Luther and M.E. Irwin (eds.) 2005. Globalizing Integrated Pest Management: A participatory research process. Blackwell Publishing, Ames Iowa. 338 pp.

Rajotte, E.G., G. W. Norton, G.C. Luther, V. Barrera and K.L. Heong. IPM Transfer and Adoption. P 143-158. In Norton, G. W., E.A. Heinrichs, G.C. Luther and M.E. Irwin (eds.) 2005. Globalizing Integrated Pest Management: A participatory research process. Blackwell Publishing, Ames Iowa. 338 pp.

Arida, G.S., B.S. Punzal, C.C. Ravina, V.P. Gapud, E.G. Rajotte and N. S. Talekar. 2004. Monitoring adult populations fo two insect pests with sex pheromone traps for effective timing of interventions agains the defoliators of onion (Allium cepa L.) grown after rice (Oryza satival.). Philippine Entomologist 18(2) 139-150.

Rajotte, E.G. and A.N.M. Rezaul Karim. 2004. Overview of the South Asia Site in Bangladesh. In Integrated Pest Management Collaborative Research Support Program. Annual Highlights (2003-2004). Pp 37-42.

Arida, G.S., C.C. Ravina, B.S. Punzal, E.G. Rajotte and N.S. Talekar. 2004. Response of the onion (Allium cepa L.) plant to simulated pest damage. Philippine Entomologist 18(2) 151-162.

Moore, K.M., E.G. Rajotte, C.W. Pitts, I. Dolinniy and O. Cholovska. 2003. Extension services in the transition from post-communist agrarian systems: The case of the plant protection stations in Ukraine. Southern Rural Sociol. 19 (1) pp. 138-165.

Arida, G. S., B. S. Punzal, C. C. Ravina Jr., V. P. Gapud, E. G. Rajotte, and N. S. Talekar. 2002. Sex pheromones in pest management monitoring adult densities of Spodoptera litura (F.) S. exigua (Hubner) (Lepidoptera: Noctuidae) in rice onion cropping systems. Philippines Rice Technical Bulletin 6(2): 36-40.  

Weiss, R. and E.G. Rajotte. 2001. IPM, Science and Science Education: The Best of Three Worlds. A Amer. Entomol. 47(2)74-77.


Research Interests

Ecological Applications

IPM, biology of fruit pests, pollination and apiculture

Pollinator Biology, Health and Ecology
Honey Bee and Pollinator Research

Dr. Rajotte is interested in crop pollination services  provided by Apis and non-Apis bees, especially as part of an integrated pest management (IPM) program. In addition, he is part of team studying the epidemiology of bee viruses.

Integrated Pest Management

Domestic and international agricultural IPM; Urban IPM implementation; IPM policy and impacts